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The course of the Mole within the airport perimeter has been altered several times since commercial flights began in 1945; however the meanders visible on the 1839 tithe map in the 1.5 km stretch immediately north of the runway were reinstated in 1999, in a £1.2 million project to facilitate airport expansion.The Mole enters Surrey to the south of Horley, where it meets the Gatwick Stream, a tributary draining Worth Forest to the southeast of Crawley.Water is able to flow out of the river through swallow holes in the bed and banks, decreasing the volume of water carried in the main channel.
In response to heavy flooding of East Molesey and Thames Ditton in September 1968, the river was modified downstream of Albany Bridge to the Thames and new flood defences were constructed.
There is only one aquifer in the drainage basin, at Fetcham, which means that the majority of the water in the river is from surface drainage, particularly from Gatwick Airport and the urban areas of Horley and Crawley, and that the flow rate responds rapidly to rainfall.
River Mole rises in Baldhorns Copse 0.7 km (0.4 mi) to the south of the village of Rusper in West Sussex.
The second-largest Sewage Treatment Works (STW) in the Mole catchment is located on the Gatwick Stream 3 km (1.9 mi) upstream of the confluence with the Mole: Crawley STW discharges 15 megalitres (15,000 m The Earlswood Brook, a tributary draining the urban area of Reigate and Redhill, joins the Mole at Sidlow.
The largest STW in the Mole catchment (Reigate STW) discharges up to 118.5 megalitres per day into the Earlswood Brook.
The Mole skirts the northern suburbs of Crawley where it is joined by its first major tributary, Ifield Brook, which drains Ifield Mill Pond.